Jonathan Balcombe’s most recent book is the New York Times bestseller What a Fish Knows: The Inner Lives of our Underwater Cousins, an extraordinary journey underwater that reveals the vast capabilities of fishes. He is also the author of the books The Exultant Ark, Second Nature, Pleasurable Kingdom, and The Use of Animals in Higher Education. 

Jonathan has three biology degrees, including a PhD in ethology (the study of animal behavior) from the University of Tennessee, and has published more than 50 scientific papers on animal behavior and animal protection. Formerly department chair for Animal Studies with the Humane Society University and senior research scientist with the Physicians Committee for Responsible Medicine, Jonathan is currently Director of Animal Sentience with the Humane Society Institute for Science and Policy in Washington, DC.